OBSERVED TIMING

1 · Calcium-linked signal

After electrical stimulation, contraction began near the peak of aequorin light—up to about 25 milliseconds after stimulation started.

calcium-sensitive lightstimuluslight peak + onsettime →≤25 ms
MINIMAL MODEL

2 · Traveling actuation

A traveling activation shortens local preferred lengths in sequence. Elasticity transmits force; environmental drag resists the change.

activation travelsdrag opposes rapid shortening
OBSERVED STRUCTURE · PROPOSED ROLE

3 · Internal damping

Vacuoles deformed by as much as 40% beside a continuous RER network. Linked-particle simulations suggest the arrangement can resist squeezing.

RER lies about 30–70 nm from vacuoles in relaxed specimensschematic cross-section · not to scale
Evidence boundary: the diagrams connect separate studies; they do not claim that the molecular force generator or the cell’s complete braking mechanism has been identified.